Fakarava, black tip reef shark and marbled groupers gathering in the pass before spawning

marbled grouper black tip reef shark reproduction Dermatolepis inermis Serranidae gathering fishes schooling fishes spawning coral pass tetamanu fakarava french polynesia loche marbrée mérou requin bordé oihe fraie rassemblement banc de poissons corail passe polynésie atoll tuamotu